Home » Publications


HEE Data Reports
  1. HEE Species List 2006-2015 (PDF: 203 KB)
  2. The Hardwood Ecosystem Experiment: Threatened and Endangered Species   (PDF: 335 KB)
Peer-Reviewed Extension Publications
  1. The Great Clearcut Controversy . 2017. (PDF: 3198 KB)
  2. The Hardwood Ecosystem Experiment: Indiana Forestry and Wildlife. 2015. (PDF: 4.9 MB)
  3. Breeding Birds and Forest Management: the Hardwood Ecosystem Experiment and the Central Hardwoods Region. 2015. (PDF: 3.2 MB)
  4. Forest Management for Reptiles and Amphibians. 2013. (PDF: 2 MB) 
Published Proceedings and Symposia
  1. Swihart, R.K., M.R. Saunders, R.A. Kalb, and C.H. Michler, eds. 2013. The Hardwood Ecosystem Experiment: a framework for studying responses to forest management. Gen. Tech. Rep. NRS-P-108. Newton Square, PA: U.S. Department of Agriculture, Forest Service, Northern Research Station. [CD ROM] 350 p.
  2. Sustainable Forestry White Paper (PDF: 17 KB)
  3. Abstracts: Managing wildlife for sustainable forests (PDF: 1.43 MB)
Peer-Reviewed Research Articles
  1. Hale, V., B. MacGowan, L. Corriveau, D. Huse, A.F. Currylow, and S. Thompson. 2017. Radio transmitter implantation and movement in the wild Timber Rattlesnake (Crotalus horridus). Journal of Wildlife Diseases, doi: 10.7589/2016-08-186
  2. Kellner, K.F. and R.K. Swihart. 2017. Herbivory on planted oak seedlings across a habitat edge created by timber harvest. Plant Ecology, 218:213-223.
  3. Kellner, K.F. and R.K. Swihart. 2017. Simulation of oak early life history and interactions with disturbance via an individual-based model, SOEL. PLoS ONE 12(6): e0179643. https://doi.org/10.1371/journal.pone.0179643
  4. MacGowan, B.J., A.F.T.Currylow and J.E.MacNeil. 2017. Short-term responses of Timber Rattlesnakes (Crotalus horridus) to even-aged timber harvests in Indiana. Forest Ecology and Management, 376: 30-36.
  5. Murray, B.D., J.D. Holand, K.S. Summerville, J.B. Dunning, M.R. Saunders, and M.A. Jenkins. 2017. Functional Diversity response to hardwood forest management varies across taxa and spatial scales. Ecological Applications 1002/eap.1532
  6. Nemes, C.E. and K. Islam. 2017. Breeding Season Microhabitat use by Cerulean Warbler (Setophaga cerulea) in an experimentally-managed forest. Forest Ecology and Management http://dx.doi.org/10.1016/j.foreco.2016.11.008
  7. Olson, M.G. and M.R. Saunders. 2017. Long-term research on managed hardwood forests in eastern North America. Forest Ecology and Management, 387: 1-2.
  8. Summerville, K.S. and R.J. Marquis. 2017. Comparing the responses of larval and adult lepidopteran communities to timber harvest using long-term, landscape-scale studies in oak-hickory forests. Forest Ecology and Management http://dx.doi.org/10.1016/j.foreco.2016.08.050
  9. Auer, S.A., K. Islam, J.R. Wagner, K.S. Summerville, and K.W. Barnes. 2016. The diet of Cerulean Warbler (Setophaga cerulea) nestlings and adult nest provisioning behaviors in S. Indiana. The Wilson Journal of Ornithology, 128(3):573-583.
  10. Barnes, K.W., K.Islam and S.A.Auer. 2016. Integrating LIDAR-derived canopy structure into cerulean warbler habitat models. Journal of Wildlife Management, 80(1): 101-116.
  11. Kellner, K.F. and R.K.Swihart. 2016. Timber harvest and drought interact to impact oak seedling growth and survival in the Central Hardwood Forest. Ecosphere 7(10): e01473
  12. Kellner, K.F., N.I.Lichti and R.K.Swihart. 2016. Midstory removal reduces effectiveness of oak (Quercus) acorn dispersal by small mammals in the Central Hardwood Forest region. Forest Ecology and Management, 375:182-190.
  13. Kellner, K.F., P.J.Ruhl, J.B.Dunning, J.K.Riegel and R.K.Swihart. 2016. Multi-scale responses of breeding birds to experimental forest management in Indiana. Forest Ecology and Management, 382:64-75.
  14. Milne, M.A., B. Foster, J.J. Lewis, L. Bishop, A. Hoffman, T. Ploss, and B. Deno. 2016. Spiders in Indiana: seventy-one new and updated distribution records. Proceedings of the Indiana Academy of Science, 125(1): 75-85.
  15. Murray, B.D., C.R.Webster, M.A.Jenkins, M.R.Saunders and G.S.Haulton. 2016. Ungulate impacts on herbaceous-layer plant communities in even-aged and uneven-aged forests. Ecosphere, 7(6): e01378
  16. Leonard, O.D., J.W.Moore, J.K.Riegel, A.R.Meier, J.B.Dunning and K.F.Kellner, R.K.Swihart. 2015. Effect of variation in forest harvest intensity on winter occupancy of barred owls and eastern screech-owls in deciduous forests of the east-central United States. Journal of Field Ornithology, 86(2): 115-129.
  17. Nemes, C.E., K.Islam and D.M.Pirtle. 2015. First documentation of a “double-decker” Cerulean Warbler (Setophaga cerulea) nest. The Wilson Journal of Ornithology, 127(3):534-538.
  18. Olson, Z.H., B.J.MacGowan, M.T.Hamilton, A.F.T.Currylow and R.N.Williams. 2015. Survival of timber rattlesnakes: Investigating individual, environmental, and ecological effects. Herpetologica, 71:274-279.
  19. Powell, G., R.M.Brattain and J.D.Holland. 2015. New distributional records for the genus Pytho. The Coleopterists’ Bulletin, 66:761-765.
  20. Ruhl, P.J., Jeffrey K. Riegel, and John B. Dunning. 2015. Confirmation of successful chestnut-sided warbler breeding in south-central Indiana. Proceedings of the Indiana Academy of Science, 124:38-42.
  21. Ruhl, P.J. and J.B.Dunning. 2015. Morphometrics of Worm-eating Warblers in south-central Indiana: hatching year and after hatching year comparison. North American Bird Bander, 43:81-84.
  22. Summerville, K.S. 2015. Species traits fail to consistently predict moth persistence in managed forest stands. Great Lakes Entomologist, 48:114-124.
  23. Wagner, J.R., K. Islam, and K. Summerville. 2015. Cerulean Warbler territory size is influenced by prey-rich tree abundance. Proceedings of the Indiana Academy of Science,124:70-79.
  24. Currylow, A.F., A.J.Johnson and R.N.Williams. 2014. Evidence of ranavirus infections among sympatric larval amphibians and box turtles. Journal of Herpetology, 48(1): 117-121.
  25. Kellner, K.F. and R.K.Swihart. 2014. Changes in small mammal microhabitat use following silvicultural disturbance. American Midland Naturalist, 172: 349-359.
  26. Kellner, K.F., J.R. Riegel and R.K. Swihart. 2014. Effects of silvicultural disturbance on acorn infestation and removal in the central hardwood forest region. New Forests, 45(2): 265-281.
  27. Kimble, S.J.A., O.E. Rhodes, Jr. and R.N. Williams. 2014. Relatedness and other finescale genetic processes in the box turtle. Conservation Genetics, 15(4): 967-979.
  28. Kimble, S.J.A., O.E.Rhodes, Jr. and R.N.Williams. 2014. Unexpectedly low rangewide population genetic structure of the imperiled eastern box turtles. . PLoS ONE 9:e92274
  29. LaGrange, S.M., S.J.A.Kimble, B.J.MacGowan and R.N.Williams. 2014. Seasonal variance in hematology and plasma chemistry values of the timber rattlesnake (Crotalus horridus). Jounal of Wildlife Diseases DOI 10.7589/2013-10-267.
  30. MacNeil, J.E. and R.N.Williams. 2014. Effects of timber harvests and silvicultural edges on terrestrial salamanders. PLoS ONE, 9(12):e114683
  31. Owen, D.A.S., E.T.Carter, M.L.Holding, K.Islam and I.T.Moore. 2014. Roads are associated with a blunted stress response in a North American pit viper. General and Comparative Endocrinology, 202: 87-92.
  32. Summerville, K.S. 2014. Do seasonal temperatures, species traits and nearby timber harvest predict variation in moth species richness and abundance in unlogged deciduous forests? Agricultural and Forest Entomology, 16: 80-86.
  33. Wagner, J.R. and K.Islam. 2014. Nest-site selection and breeding ecology of the Cerulean Warbler in southern Indiana. Northeastern Naturalist, 21: 515-528.
  34. Auer, S.A., K.Islam, K.W.Barnes and J.A.Brown. 2013. Documentation of Red-bellied Woodpecker predation of a Cerulean Warbler nestling. The Wilson Journal of Ornithology, 125(3) 642-646.
  35. Currylow, A.F., M.S.Tift, J.L.Meyer, D.E.Crocker and R.N.Williams. 2013. Seasonal variations in plasma vitellogenin and sex steroids in male and female eastern box turtles (Terrapene c. carolina). General and Comparative Endocrinology, 180:48-55.
  36. Kaminski, K.J. and K. Islam. 2013. Effects of forest treatments on abundance and spatial characteristics of cerulean warbler territories. American Midland Naturalist, 170:111 – 120.
  37. Kellner, K.F., N.A.Urban and R.K.Swihart. 2013. Short-term responses of small mammals to timber harvest in the Central Hardwoods. Journal of Wildlife Management, 77(8): 1650-1663.
  38. MacNeil, J.E. and R.N.Williams. 2013. Effectiveness of two artificial cover objects in sampling terrestrial salamanders. Herpetological Conservation and Biology, 8(3): 552-560.
  39. Summerville, K.S. 2013. Forest lepidopteran communities are more resilient to shelterwood harvest compared to more intensive logging regimes. Ecological Applications, 23(5): 1101-1112.
  40. Summerville, K.S., J.L. Lane and D. Courard-Hauri. 2013. Stability in forest lepidopteran communities: how sensitive are pest species to experimental forest management. Insect Conservation and Diversity, 6:(3)265-275.
  41. Currylow, A.F., B.J.MacGowan and R.N.Williams. 2012. Hibernal thermal ecology of eastern box turtles within a managed forest landscape. Journal of Wildlife Management, 77(2): 326-335.
  42. Currylow, A.F., B.J.MacGowan and R.N.Williams. 2012. Short-term forest management effects on a long-lived ecotherm. PLoS ONE, 7(7):e40473.
  43. Islam, K., J.Wagner, R.Dibala, M.M.MacNeil, K.J.Kaminski and L.P.Young. 2012. Cerulean Warbler (Setophaga cerulea) response to changes in forest structure in Indiana. Ornitologia Neotropical, 23: 335-341
  44. Kimble, S.J.A. and R.N.Williams. 2012. Temporal variance in hematologic and plasma bi0chemical reference intervals for free-ranging eastern box turtles (Terrapene carolina carolina). Journal of Wildlife Diseases, 48(3):799-802.
  45. Urban, N.A., R.K.Swihart, M.C.Malloy and J.B.Dunning. 2012. Improving selection of indicator species when detection is imperfect. Ecological Indicators, 15: 188-197.
  46. Currylow, A.F., P.A.Zollner, B.J.MacGowan and R.N.Williams. 2011. A survival estimate of Midwestern adult Eastern box turtles using radio telemetry. The American Midland Naturalist, 165(1): 143-149.
  47. Summerville, K.S. 2011. Managing the forest for more than the trees: effects of experimental timber harvest on forest Lepidoptera. Ecological Applications, 21(3): 806-816.
  48. Urban, N.A. and R.K.Swihart. 2011. Small mammal responses to forest management for oak regeneration in southern Indiana. Forest Ecology and Management, 261:353-361.
  49. Holland, J.D. 2010. Isolating spatial effects on beta diversity to inform forest landscape planning. Landscape Ecology, 25(9): 1349-1362.
  50. Summerville, K.S., D.Courard-Hauri and M.M.Dupont. 2009. The legacy of timber harvest: Do patterns of species dominance suggest recovery of Lepidopteran communities in managed hardwood stands? Forest Ecology and Management, 259(1): 8-13.
  51. Summerville, K.S., M.M.Dupont, A.V.Johnson and R.L.Krehbiel. 2008. Spatial structure of forest Lepidopteran communities in oak hickory forests of Indiana. Environmental Entomology, 37(5): 1224-1230.
Published Theses and Dissertations
  1. Nelson, D. 2017. Demographic responses of small mammals to disturbance induced by forest management. M.S. Thesis. Purdue University, West Lafayette, Indiana.
  2. Nemes, C.E. 2016. Cerulean Warbler (Setophaga cerulea) microhabitat use in southern Indiana. M.S. Thesis. Ball State University, Muncie, Indiana
  3. Nolder, A.D. 2016. Bat species activity related to silvicultural treatments in hardwood forests managed for oak regeneration, with an examination of the influence of microphone height on acoustic survey results. M.S. Thesis. Indiana University of Pennsylvania, Indiana, Pennsylvania.
  4. Caldwell, K.L. 2015. A comparison of bat activity in a managed central hardwood forest. M.S. Thesis. Ball State University, Muncie, Indiana
  5. Kellner, K.F. 2015. Interactive effects of animals and silviculture on the early life history of oak. Ph.D. Dissertation. Purdue University, West Lafayette, IN.
  6. Auer, S. 2014. Cerulean Warbler (Setophaga cerulea) nest provisioning behaviors and the effects of forest management practices on nest success in southern Indiana. M.S. Thesis. Ball State University, Muncie, Indiana.
  7. Badin, H.A. 2014. Habitat selection and roosting ranges of northern long-eared bats (Myotis septentrionalis) in an experimental hardwood forest system. M.S. Thesis. Ball State University, Muncie, Indiana
  8. Barnes, K. 2014. Assessing the influence of social information and habitat structure on Cerulean Warbler breeding distribution in southern Indiana. M.S. Thesis. Ball State University, Muncie, Indiana.
  9. Dibala, R. 2012. Population response of a declining songbird to silviculture: how Cerulean Warbler (Setophaga cerulea) territory size and settlement patterns fare in the face of forest disturbance. M.S. Thesis. Ball State University, Muncie, IN.
  10. Kellner, K.F. 2012. Temporal dynamics of mast and small mammals: short-term responses to silviculture. M.S. Thesis. Purdue University, West Lafayette, Indiana.
  11. Kimble, S.J.A. 2012. Population genetics of the eastern box turtle, Terrapene c. carolina. Ph.D. Dissertation. Purdue University, West Lafayette, Indiana
  12. Malloy, M.C. 2012. Effects of forest management on breeding bird populations on mixed deciduous forests of southern Indiana. M.S. Thesis. Purdue University, West Lafayette, Indiana.
  13. Meier, A.R. 2012. Aspects of epicormic development in Quercus alba (L.) and other eastern North American oak species in relation to genetics, tree vigor and silvicultural treatments. M.S. Thesis. Purdue University, West Lafayette, Indiana.
  14. Wagner, J. 2012. Cerulean Warbler population and breeding response to recent silviculture and influences of prey availability on avian nesting ecology. M.S. Thesis. Ball State University, Muncie, Indiana.
  15. Caylor, M.K. 2011. Impacts of different forest tree-harvest methods on diets and populations of insectivorous forest bats. . Master of Science Thesis. Indiana State University, Terre Haute, IN.
  16. Currylow, A.F. 2011. Effects of forest management on the ecology and behavior of Eastern box turtles. M.S. Thesis. Purdue University, West Lafayette, Indiana.
  17. MacNeil, J.E. 2011. The effects of forest management on terrestrial salamanders. M.S. Thesis. Purdue University, West Lafayette, Indiana.
  18. Kaminski, K.J. 2010. Cerulean Warbler Initial Response to Silviculture Treatments in Southern Indiana. M.S. Thesis. Ball State University, Muncie, Indiana.
  19. MacNeil, M.M. 2010. Does timber harvesting affect cerulean warbler foraging ecology? M.S. Thesis. Ball State University, Muncie, Indiana.
  20. Sheets, J.J. 2010. Impact of forest management techniques on bats with a focus on the endangered Indiana myotis (Myotis sodalis). M.S. Thesis. Indiana State University, Terre Haute, Indiana.
  21. Urban, N.A. 2010. Improving inferences on wildlife responses to oak-centered forest management with models that account for imperfect detection. M.S. Thesis. Purdue University, West Lafayette, Indiana.
  22. Rogers, S.C. 2008. Public acceptability of management practices on the Morgan-Monroe State Forest: Examining the influence of information, values, and visual characteristics. M.S. Thesis. Purdue University, West Lafayette, Indiana.
  23. Young, L.P. 2008. Distribution and foraging ecology of Cerulean Warblers in southern Indiana. M.S. Thesis. Ball State University, Muncie, Indiana.